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NAES header image
NAES header image


Kelley Stewart

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Associate Professor

Department of Natural Resources and Environmental Science
University of Nevada/Mail Stop 186
1664 North Virginia Street
Reno,  Nevada   89557

Office: (775) 784-4314
Fax: 784-4583

Building: Max Fleischmann Agriculture,  Office 220F
Personal Web:

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BS (1993) University of California Davis
MS (1997) Caesar Kleberg Wildlife Research Institute Texas A&M University - Kingsville
PhD (2004) Dept of Biology & Wildlife, & Institute of Arctic Biology, University of Alaska Fairbanks


I am interested in population ecology of large mammals, and effects of density dependence on population growth, movement patterns, habitat selection, and both inter- and intraspecific competition.  I am also interested in community-level effects of population dynamics of large mammals, such as changes in plant growth, community structure, and trophic cascades.


  • 2012 Teacher of the Year Award College of Agriculture, Biotechnology and Natural Resources
  • 2005 Post-Doctoral Fellowship (2 years) Dept. Biological Sciences, Idaho State University
  • 2004 Graduate Fellowship (6 months): Rob and Bessie Welder Wildlife Foundation
  • 2003 Graduate Fellowship (1 year): Rob and Bessie Welder Wildlife Foundation
  • 2002 Robert I. Larus Prize: 53rd Arctic American Association for Advancement of Science Conference (Best presentation by a graduate student)
  • 2002 Graduate Fellowship (6 months): Rob and Bessie Welder Wildlife Foundation
  • 2001 Graduate Fellowship: Graduate School, University of Alaska Fairbanks
  • 1996 Gamma Sigma Delta, Agricultural Honor Society
  • 1994 Graduate Fellowship (3 years): Rob and Bessie Welder Wildlife Foundation


  • Chair Technical Session, Nevada Chapter of The Wildlife Society Symposium, Reno, NV. January 2012
  • Chair Honorary Membership and Special Recognition Service Award, The Wildlife Society, 2011 to present
  • Presentation for Nevada Board of Wildlife Commissioners, March 2011
  • Chair Technical Session, Nevada Chapter of The Wildlife Society Symposium, Reno, NV. January 2011
  • Chair Technical Session North American Moose Conference and Workshop Pocatello, ID – 2009    
  • Faculty advisor: UNR Student Chapter of the Wildlife Society 2008 - Present
  • Resolutions Committee: American Society of Mammalogists 2004 - 2010
  • Organizing Committee: 2009 Annual Meeting of The Wildlife Society
  • Organizing Committee: 10th Western Black Bear Workshop 2009


Simpson, N. O., Stewart, K., Schroeder, C., Cox, M., Huebner, K., Wasley, T. 2016, Overpasses and underpasses: effectiveness of crossing structures for migratory ungulates., Journal of Wildlife Management, 80(8), 1370-1378.   Read More...
Downs, CJ and KM Stewart. 2015, A primer in ecoimmunology and immunology for wildlife research and management., California Fish and Game 100 (3):371-395  
Blum, M., Stewart, K., Schroeder, C. 2015, Effects of large scale gold mining on migratory behavior of a large herbivore., Ecosphere   Read More...
Downs, C. J., Stewart, K., Dick, B. L. 2015, Investment in Constitutive Immune Function by North American elk experimentally maintained at two different popualtion densities., Plos One, 10(5), 17.  
Stewart, K., Walsh, D. R., Kie, J. G., Dick, B. L., Bowyer, R. T. 2015, Sexual Segregation in North American elk: the role of density dependence., Ecology and Evolution   Read More...
McKee, C., Stewart, K., Sedinger, J. S., Bush, A., Darby, N., Hughson, D., Bleich, V. 2015, Spatial distribution and resource selection by mule deer in an arid environment: responses to provision of water., Journal of Arid Environments, 122, 76-84.  
Downs, C. J., Stewart, K. 2014, A primer in ecoimmunology and immunology for wildlife research and management., California Fish and Game, 100(3), 369-393.  
Bowyer, R. T., Bleich, V. C., Stewart, K., Whiting, J. C., Monteith, K. L. 2014, Density Dependence in Ungulates: a Review of Causes, and Concepts with Some Clarification., California Fish and Game, 100(3), 550-572.  
Andreasen, A. M., Stewart, K., Longland, W. S., Beckman, J. P., Forister, M. L. 2012, Identification of source-sink dynamics in mountain lions of the Great Basin., Molecular Ecology/Blackwell, 21(23), 5689–5701.  
Morano, S., Stewart, K., Sedinger, J. S., Nicolai, C., Vavra, M. 2012, Life-history strategies of North American elk: trade-offs associated with reproduction and survival., Journal of Mammalogy, 93(6), 11.  
Kelley M. Stewart, R. Terry Bowyer, Brian L. Dick & John G. Kie 2011, Effects of density dependence on diet composition of North American elk Cervus elaphus and mule deer Odocoileus hemionus: an experimental manipulation, Wildl. Biol. 17: 417-430   Read More...
Stewart, K., Bowyer, R. T., Dick, B. L., Kie, J. G. 2011, Population density of North American elk: density-dependent effects on diet composition of mule deer and elk, Willdife Biology, 17(4), 417-430   Read More...
Bowyer, R. T., Rachlow, J. L., Stewart, K., VanBallenberghe, V. 2011, Vocalizations by alaskan moose: female incitation of male aggression, Behavioral Ecology and Sociobiology, 65(12), 2251-2260   Read More...
Nova O. Simpson, Kelley M. Stewart, And Vernon C. Bleich 2011, What have we learned about water develpments for wildlife? Not enough!, California Fish and Game 97(4):190-209   Read More...
Stewart, K. M., R. T. Bowyer, J. G. Kie, and M. A. Hurley. 2010, Spatial distributions of mule deer and North American elk: resource partitioning in a sage-steppe environment., American Midland Naturalist 163:400-412   Read More...
Stewart, K. M., R. T. Bowyer, J. G. Kie, B. L. Dick, and R. W. Ruess. 2009, Population density of North American elk: effects on plant diversity., Oecologia 161:303-312   Read More...
Whiting, J. C., K. M. Stewart, R. T. owyer, and J. T. Flinders. 2009, Reintroduced bighorn sheep: do felames adjust maternal care to compensate for late-born young?, European Journal of Wildlfie Research. Published online November 2009.   Read More...
K.M. Stewart, R. T. Bowyer, R.W. Ruess, B.L. Dick, J. G. Kie 2007, Herbivore optimization by North American elk: consequences for theory and management, Wildlife Monographs 167: 1–24   Read More...
Stewart, K. M., R. T. Bowyer, B. L. Dick, B. K. Johnson, and J. G. Kie. 2006, Density Dependence in North American Elk: An Experimental Test, Oecologia 143:85-93   Read More...
Kelley M. Stewart, R. Terry. Bowyer, Brian L. Dick, Bruce K. Johnson, John G. Kie 2005, Density-dependent effects on physical condition and reproduction in North American elk: an experimental test, Oecologia (2005) 143: 85–93   Read More...
Stewart, K. M., R. T. Bowyer, R. W. Ruess, B. L. Dick, and J. G. Kie. 2005, Herbivore optimization in North American elk: consequences for theory and management., Wildlife Monographs 167   Read More...
Stewart, K. M., R. T. Bowyer, J. G. Kie, B. L. Dick, and M. Ben-David 2003, Niche partitioning among mule deer, elk, and cattle: do stable isotopes reflect dietary niche?, Écoscience 10:297-302   Read More...
Stewart, K. M., T. E. Fulbright, D. L. Drawe, and R. T. Bowyer. 2003, Sexual segregation in white-tailed deer: responses to habitat manipulations, Wildlife Society Bulletin 31:1210-1217   Read More...
R. Terry Bowyer, Kelley M. Stewart, Scott A. Wolfe, Gail M. Blundell, Karin L Lehmkuhl, Philip J. Joy, Thomas J. Mcdonough, John G. Kie 2002, Assessing Sexual Segregation In Deer, Journal Of Wildlife Management 66(2):536-544   Read More...
R. Terry Bowyer, Kelley M. Stewart, Becky M. Pierce, Kris J. Hundertmark, and William C. Gasaway 2002, Geographical Variation In Antler Morphology Of Alaskan Moose: Putative Effects Of Habitat And Genetics, ALCES VOL. 38: 155-165   Read More...
Stewart, K. M., J. G. Kie, R. T. Bowyer, N. J. Cimon, and B. K. Johnson. 2002, Temporospatial distribution of large herbivores: resource partitioning and competitive displacement., Journal of Mammalogy 83:229-244   Read More...
Kelley M. Stewart, R. Terry Bowyer,* John G. Kie, Norman J. Cimon, And Bruce K. Johnson 2002, Temporospatial Distributions Of Elk, Mule Deer, And Cattle: Resource Partitioning And Competitive Displacement, Journal of Mammalogy, 83(1):229–244, 2002   Read More...
R. Terry Bowyer,* Kelley M. Stewart, John G. Kie, And William C. Gasaway 2001, Fluctuating Asymmetry In Antlers Of Alaskan Moose: Size Matters, Journal of Mammalogy, 82(3):814–824   Read More...
Kelley M. Stewart, R. Terry Bowyer, John G. Kie, and William C. Gasaway 2000, Antler Size Relative To Body Mass In Moose: Tradeoffs Associated With Reproduction, ALCES VOL. 36: 77-83   Read More...
K.M. Stewart, T.E. Fulbright, D.L. Drawe 2000, White-Tailed Deer Use Of Clearings Relative To Forage Availability, J. Wildl. Manage. 64(3):2000   Read More...
Book or Chapter(s) in Books
Bowyer, R. T., Stewart, K., Long, R., Monteith, K. L. 2012, Wildlife Ecology, In A.H. El-Shaarawi, and W.W. Piegorsch (Ed.), Encyclopedia of Environmetrics (vol. 2nd edition, pp. 2963-2968). New York, New York: John Wiley and Sons, LTd.  
Stewart, K., Bowyer, R. T., Weisberg, P. J. 2011, Spatial Use of Landscapes by White-Tailed Deer., In David Hewitt (Ed.), Biology and Management of White-Tailed Deer.   Read More...
John G. Kie, R. Terry Bowyer And Kelley M. Stewart 2003, Ungulates in western coniferous forests: habitat relationships, population dynamics, and ecosystem processes, Management and Conservation in the Coniferous Forests of Western North America, Edited by Cynthia J. Zabel, Cambridge University Press   Read More...
News Letters
Stewart, K. 2013, Use of Artificial Water Sources by Mule Deer in Mojave National Preserve, In Debra Hughson, Science Advisor, NPS (Ed.), (vol. May 2013, pp. 4). Barstow, CA: Mojave National Preserve Science Newsletter.  
Technical Publication
Stewart, K., Simpson, N. O., Tian, Z., Attah, I., Cox, M., Wasley, T. 2015, Effectiveness of wildlife crossing structures for mule deer and other wildlife in Nevada, (vol. 101-10-803, pp. 29). Carson City, Nevada: Nevada Department of Transportation.  

Highways and Interstate freeways not only fragment movement corridors for wildlife, but result in property damage, injuries and sometimes fatalities for humans and increased mortality of wildlife from collisions with vehicles. Crossing structures for wildlife are an important tool to allow safe passage for wildlife across roadways. We documented use by mule deer (Odocoileus hemionus) of overpasses and underpasses at 2 sites in eastern Nevada and evaluated their success for removing mule deer from U.S. Highway 93 (US 93). Our metrics to evaluate success included the number of animals that crossed each structure, passage rate (percentage of approaches that resulted in successful crossings) at each structure, and mortalities of mule deer on US 93 during migrations. We used a benefit-cost analysis of the wildlife overpass at 10-Mile Summit to determine the economic benefit of the structures. Crossing structures were immediately successful and mortalities declined with each subsequent migration although the number of deer that crossed the structures increased. Most of the deer that crossed US 93 crossed the overpass at 10-Mile Summit. Over the study period the total number of crossings increased at the overpasses and declined at the underpasses at both sites, shifting to more use of the overpasses. Passage rates were consistently greater at the overpasses across all migrations, regardless of site. Both types of structures proved to be successful for removing migratory ungulates from the roadway, and the benefit-cost analysis indicated that construction and maintenance of the overpass was beneficial. Therefore, the US 93 crossing structures succeeded in making roadways safer for both wildlife and motorists.